Harpalinae

















Harpalinae

Harpalini.jpg
Undetermined Harpalinae species

Scientific classification
Kingdom:

Animalia
Phylum:

Arthropoda
Class:

Insecta
Order:

Coleoptera
Suborder:

Adephaga
Family:

Carabidae
Subfamily:

Harpalinae

Bonelli, 1810

Supertribes

  • Anthiitae

  • Chlaeniitae

  • Dryptitae

  • Harpalitae

  • Lebiitae

  • Pentagonicitae

  • Platynitae

  • Pterostichitae

Harpalinae is a huge subfamily of ground beetles that contains 20,000 species[citation needed]. A rarely used common name for the subfamily is the harp beetles[citation needed]. The Harpalinae contain the most apomorphic ground beetles, displaying a wide range of forms and behaviors. Some are, rare among ground beetles, omnivores or even herbivores.


Many closely related subfamilies have been treated as subordinate taxa of the Harpalinae by various authors. Among these are the Dryptinae, Lebiinae (including Cyclosominae, Mormolycinae, Odacanthinae, Perigoninae), Licininae (including Chlaeniinae, Oodinae), Orthogoniinae, Panagaeinae, Platyninae, Pseudomorphinae, Pterostichinae (including Zabrinae). Here, they are considered independent families within the harpaline (sensu lato) assemblage, and this is also tentatively assumed for the enigmatic monotypic genus Ginema.



Systematics


At least 4 large and several smaller supertribes can be distinguished. The former "Pterostichitae" on the other hand seem to be, as presently delimited, a paraphyletic assemblage, but probably also contains a number of tribes closer to each other than to other Harpalinae. The clade around the type genus Pterostichus is thus recognized as subfamily Pterostichinae here.[1]



Genera


The Harpalinae include the following genera:[2][verification needed]





Harpalus affinis




  • Acinopus Dejean, 1821


  • Acupalpus Latreille, 1829


  • Afromizonus Basilewsky, 1947


  • Agonidus Casey, 1914


  • Allendia Noonan, 1974


  • Allocinopus Broun, 1903


  • Allosiopelus Ito, 1995


  • Amblygnathus Dejean, 1829


  • Amblystomus Erichson, 1837


  • Amerinus Casey, 1884


  • Amphasia Newman, 1836


  • Angionychus Klug, 1853


  • Anisocnemus Chaudoir, 1843


  • Anisodactylus Dejean, 1829


  • Anisostichus Emden, 1953


  • Anomostomus Laferte-Senectere, 1853


  • Anthracus Motschulsky, 1850


  • Athrostictus Bates, 1878


  • Axinotoma Dejean, 1829


  • Aztecarpalus Ball, 1970


  • Barysomus Dejean, 1829


  • Batoscelis Dejean, 1836


  • Bleusea Bedel, 1897


  • Boeomimetes Peringuey, 1896


  • Bottchrus Jedlicka, 1935


  • Bradybaenus Dejean, 1829


  • Bradycellus Erichson, 1837


  • Bradycidus Casey, 1914


  • Bronislavia Semenov, 1891


  • Carenochyrus Solsky, 1874


  • Carterus Dejean, 1830


  • Cenogmus Sloane, 1898


  • Chilotomus Chaudoir, 1842


  • Chydaeus Chaudoir, 1854


  • Coleolissus Bates, 1892


  • Crasodactylus Guerin-Meneville, 1847


  • Cratacanthus Dejean, 1829


  • Cratognathus Dejean, 1829


  • Cratosoma Jeannel, 1948


  • Criniventer van Emdem, 1953


  • Cyptomicrus Vinson, 1939


  • Daptus Fischer von Waldheim, 1823


  • Diachromus Erichson, 1837


  • Dicheirotrichus Jacqelin du Val, 1857 (including Trichocellus)


  • Dicheirus Mannerheim, 1843


  • Dioryche W.S. MacLeay, 1825


  • Discoderus LeConte, 1853


  • Ditomus Bonelli, 1810


  • Dixus Billberg, 1820


  • Dregus Motscheisky, 1864


  • Ectinothorax Alluaud, 1941


  • Egadyla Alluaud, 1916


  • Eocarterus Stichel, 1923


  • Eriophonus Tschitscherine, 1901


  • Eucarterus Reitter, 1900


  • Euryderus LeConte, 1846


  • Euthenarus Bates, 1874


  • Gaioxenus Broun 1910


  • Geodromus Dejean, 1829


  • Geopinus LeConte, 1848


  • Gnathaphanus W.S. MacLeay, 1825


  • Goniocellus Casey, 1914


  • Graniger Motschulsky, 1864


  • Gugheorites Basilewsky, 1951


  • Gynandromorphus Dejean, 1829


  • Hakaharpalus Larochelle & Lariviere, 2005


  • Haplaner Chaudoir, 1878


  • Haplanister Moore, 1996


  • Harpalinus Jeannel, 1946


  • Harpalobrachys Tschitscherine, 1899


  • Harpalodiodes Bousquet, 2002


  • Harpalomimetes Schauberger, 1933


  • Harpalomorphus Peringuey, 1896


  • Harpalus Latreille, 1802 (including Harpalobrachys)


  • Harpathaumas Basilewsky, 1947


  • Harponixus Basilewsky, 1950


  • Hartonymus Casey, 1914


  • Hemiaulax Bates, 1892


  • Heteracantha Brulle, 1834


  • Hippoloetis Laporte, 1835


  • Hyphaereon W.S. MacLeay, 1825


  • Hypharpax W.S. McLeay, 1825


  • Idiomelas Tschitscherine, 1900


  • Indiophonus N.Ito, 1996


  • Kaffovatus Clarke, 1972


  • Kenyacus Alluaud, 1917


  • Kiwiharpalus Larochelle & Lariviere, 2005


  • Kupeharpalus Larochelle & Lariviere, 2005


  • Lecanomerus Chaudoir, 1850


  • Liochirus Tschitscherine, 1897


  • Liodaptus Bates, 1889


  • Lioholus Tschitscherine, 1897


  • Loxoncus Schmidt-Gobel, 1846


  • Machozetus Chaudoir, 1850


  • Maoriharpalus Larochelle & Lariviere, 2005


  • Meroctenus Gemminger et Harold, 1868


  • Microderes Faldermann, 1836


  • Nemaglossa Solier, 1849


  • Neoaulacoryssus Noonan, 1985


  • Neodiachipteryx Noonan, 1985


  • Neohyparpalus Clarke, 1981


  • Neophygas Noonan, 1976


  • Nesarpalus Bedel, 1897


  • Nipponoharpalus Habu, 1973


  • Nornalupia Kataev, 2002


  • Nothodaptus Maindron, 1906


  • Notiobia Perty, 1830


  • Notospeophonus Moore, 1962


  • Odontocarus Solier, 1835


  • Oedesis Motschulsky, 1850


  • Oesyperus Andrewes, 1923


  • Omostropus Peringuey, 1896


  • Ooidius Chaudoir, 1847


  • Oosoma Nietner, 1857


  • Ophoniscus Bates, 1892


  • Ophonus Dejean, 1821


  • Orthogenium Chaudoir, 1835


  • Oxycentrus Chaudoir, 1854


  • Pachycarus Solier, 1835


  • Pachytrachelus Chaudoir, 1852


  • Panagrius Andrewes, 1933


  • Pangus Dejean, 1821


  • Parabaris Broun, 1881


  • Paramecus Dejean, 1829


  • Parasiopelus Basilewsky, 1950


  • Paregaploa Muller, 1947


  • Parophonus Ganglbauer, [1891]


  • Paulianoscirtus Basilewsky, 1976


  • Pelmatellus Bates, 1882


  • Penthophonus Reitter, 1900


  • Penthus Chaudoir, 1843


  • Phanagnathus Basilewsky, 1950


  • Phobophorus Motschulsky, 1850


  • Pholeodytes Britton, 1962


  • Phorticosomus Shaum, 1863


  • Phyrometus Basilewsky, 1946


  • Piosoma LeConte, 1848


  • Platymetopsis Ball et Maddison, 1987


  • Platymetopus Dejean, 1829


  • Pogonodaptus Horn, 1881


  • Polpochila Solier, 1849


  • Prakasha Andrewes, 1919


  • Proditomus Schauberger, 1934


  • Progonochaetus J. Muller, 1938


  • Pseudanisotarsus Noonan, 1973


  • Pseudaristus Reitter, 1900


  • Pseudodiachipteryx Burgeon, 1936


  • Pseudognathaphanus Schauberger, 1932


  • Pseudohyparpalus Basilewsky, 1946


  • Pseudorhysopus Kataev et Wrase, 2001


  • Pseudoselenophorus Peringuey, 1896


  • Psychristus Andrewes, 1930


  • Rhabidius Basilewsky, 1948


  • Rhysopus Andrewes, 1929


  • Scybalicus Schaum, 1862


  • Selenophorus Dejean, 1831


  • Selenotichnus Kataev, 1999


  • Siopelus Murray, 1859


  • Stenolophidius Jeannel, 1948


  • Stenolophus Dejean, 1821


  • Stenomorphus Dejean, 1831


  • Syllectus Bates, 1878


  • Trachysarus Reed, 1874


  • Trichopselaphus Chaudoir, 1843


  • Trichotichnus A.Morawitz, 1863


  • Trichoxycentrus N. Ito, 2000


  • Triplosarus Bates, 1874


  • Tropicoritus Alluaud, 1917


  • Tschitscherinellus Csiki, 1906


  • Tuiharpalus Larochelle & Lariviere, 2005


  • Typsiharpalus Tschitscherine, 1901


  • Uenanthracus Kasahara, 1994


  • Veradia Castelnau, 1867


  • Xenodochus Andrewes, 1941


  • Xenophonus Muller, 1942


  • Xestonotus LeConte, 1853





References







  • .mw-parser-output .smallcapsfont-variant:small-caps
    Carabidae of the World Database (CWD) (2008): Trees of family Carabidae. Retrieved 2008-JUL-24.


  1. ^ CWD (2008)


  2. ^ "Harpalinae Bonelli, 1810". Carabidae of the World. 2012. Retrieved 11 Nov 2012..mw-parser-output cite.citationfont-style:inherit.mw-parser-output qquotes:"""""""'""'".mw-parser-output code.cs1-codecolor:inherit;background:inherit;border:inherit;padding:inherit.mw-parser-output .cs1-lock-free abackground:url("//upload.wikimedia.org/wikipedia/commons/thumb/6/65/Lock-green.svg/9px-Lock-green.svg.png")no-repeat;background-position:right .1em center.mw-parser-output .cs1-lock-limited a,.mw-parser-output .cs1-lock-registration abackground:url("//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Lock-gray-alt-2.svg/9px-Lock-gray-alt-2.svg.png")no-repeat;background-position:right .1em center.mw-parser-output .cs1-lock-subscription abackground:url("//upload.wikimedia.org/wikipedia/commons/thumb/a/aa/Lock-red-alt-2.svg/9px-Lock-red-alt-2.svg.png")no-repeat;background-position:right .1em center.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registrationcolor:#555.mw-parser-output .cs1-subscription span,.mw-parser-output .cs1-registration spanborder-bottom:1px dotted;cursor:help.mw-parser-output .cs1-hidden-errordisplay:none;font-size:100%.mw-parser-output .cs1-visible-errorfont-size:100%.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration,.mw-parser-output .cs1-formatfont-size:95%.mw-parser-output .cs1-kern-left,.mw-parser-output .cs1-kern-wl-leftpadding-left:0.2em.mw-parser-output .cs1-kern-right,.mw-parser-output .cs1-kern-wl-rightpadding-right:0.2em










-

Popular posts from this blog

How to check contact read email or not when send email to Individual?

Image
Clash Royale CLAN TAG #URR8PPP up vote 1 down vote favorite I'm using WordPress 4.9.8, CiviCRM to 5.5.1, I usually send email to contact by Search> Find contacts View contact details Action> Send email Send email ok, Contact received mail ok like picture But status only Email sent though contact read email or not. So, can CiviCRM can change status to Email read when contact read email? wordpress email share | improve this question asked Sep 26 at 0:12 ToanLuong 49 9 add a comment  |  up vote 1 down vote favorite I'm using WordPress 4.9.8, CiviCRM to 5.5.1, I usually send email to contact by Search> Find contacts View contact details Action> Send email Send email ok, Contact received mail ok like picture But status only Email sent though contact read email or not. So, can CiviCRM can change status to Email read when contact read email? wordpress email share | improve this questi...
Read more

Displaying single band from multi-band raster using QGIS

Image
Clash Royale CLAN TAG #URR8PPP 1 How can I extract a single band from multi-band raster in QGIS? I have an remote sensed image which has 6 bands (including NDVI band), I want to display each band separately, but have no idea how to do. I have seen some questions similar here but none worked for me. The original image (has 6 bands) is: I want to display the band 6 which should be like this: But I tried gdal_translate, and couldn't get the correct result. What I have got is: qgis raster multi-band share | improve this question edited Mar 5 at 0:53 Summer asked Mar 4 at 6:42 Summer Summer 23 6 Is this any help gis.stackexchange.com/questions/220658/… ? if not gis.stackexchange.com/questions/62133/… might help. – Michael Stimson Mar 4 at 6:46 Thanks for answering but when I used gdal_translate, qgis showed that 'Error 4: Kayena.tif: No such file or directory". Would you know how to fi...
Read more

How many registers does an x86_64 CPU actually have?

Image
Clash Royale CLAN TAG #URR8PPP up vote 2 down vote favorite I am currently learning reverse engineering and am studying the flags register. I had in my mind that rflags was just another name for one of the 16 general purpose registers, for example rax or rbx . But it looks like rflags is actually an additional register. So that makes 17 registers in total... how many more could there be? I have spent at least an hour on this and found numerous different answers. The best answer so far is this, which says that there are 40 registers in total. 16 General Purpose Registers 2 Status Registers 6 Code Segment Registers 16 SSE Registers 8 FPU/MMX Registers But if I add that up, I get 48. Could anybody provide an official answer on how many registers an x86_64 CPU has (e.g. an Intel i7). Additionally, I have seen references to 'hardware' and 'architectural' registers. What are those registers and how many are there? register x86-64 share | improve this...
Read more