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Acalyptratae
Marsh fly (Sciomyzidae)
Scientific classification
Kingdom:	Animalia
Phylum:	Euarthropoda
Class:	Insecta
Order:	Diptera
Section:	Schizophora
Subsection:	Acalyptratae
Superfamilies
Carnoidea Conopoidea Diopsoidea Ephydroidea Lauxanioidea Lonchaeoidea Nerioidea Opomyzoidea Sciomyzoidea Sphaeroceroidea Tephritoidea

The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, as opposed to the Calyptratae. All forms of the name refer to the lack of calypters in the members of this subsection of flies. An alternative name, Acalypterae is current, though in minority usage. It was first used by Justin Pierre Marie Macquart in 1835 for a section of his tribe Muscides; he used it to refer to all acalyptrates plus scathophagids and phorids, but excluding Conopidae.

The confusing forms of the names stem from their first usage; Acalyptratae and Acalyptrata actually are adjectival forms in New Latin. They were coined in the mid 19th century in contexts such as "Muscae Calyptratae and Acalyptratae" and "Diptera Acalyptrata", and the forms stuck.^[1]

The Acalyptratae are a large assemblage, exhibiting very diverse habits, with one notable and perhaps surprising exception: no known acalyptrates are obligate blood-feeders (hematophagous), though blood feeding at various stages of the life history is common throughout other Dipteran sections.

Classification

• Subsection Acalyptratae
  
  • Superfamily Conopoidea
    
    • Conopidae
  
  
  • Superfamily Tephritoidea
    
    • Ctenostylidae
    
    
    • Pallopteridae
    
    
    • Piophilidae
    
    
    • Platystomatidae
    
    
    • Pyrgotidae
    
    
    • Richardiidae
    
    
    • 
      Tephritidae (including Tachiniscididae)
    
    
    • Ulidiidae
    
    
  
  
  • Superfamily Nerioidea
    
    • Cypselosomatidae
    
    
    • Micropezidae
    
    
    • Neriidae
    
    
    • Pseudopomyzidae
    
    
  
  
  • Superfamily Diopsoidea
    
    • Diopsidae
    
    
    • Gobryidae
    
    
    • Megamerinidae
    
    
    • Nothybidae
    
    
    • Psilidae
    
    
    • Somatiidae
    
    
    • Syringogastridae
    
    
    • Strongylophthalmyiidae
    
    
    • Tanypezidae
    
    
  
  
  • Superfamily Sciomyzoidea
    
    • Coelopidae
    
    
    • Dryomyzidae
    
    
    • Helosciomyzidae
    
    
    • Helcomyzidae
    
    
    • Heterocheilidae
    
    
    • Ropalomeridae
    
    
    • Sepsidae
    
    
    • 
      Sciomyzidae (including Huttoninidae and Phaeomyiidae)
    
    
  
  
  • Superfamily Sphaeroceroidea
    
    • Chyromyidae
    
    
    • Heleomyzidae
    
    
    • Nannodastiidae
    
    
    • Sphaeroceridae
    
    
  
  
  • Superfamily Lauxanioidea
    
    • Celyphidae
    
    
    • Chamaemyiidae
    
    
    • Lauxaniidae
    
    
  
  
  • Superfamily Opomyzoidea
    
    • Agromyzidae
    
    
    • Anthomyzidae
    
    
    • Asteiidae
    
    
    • Aulacigastridae
    
    
    • Clusiidae
    
    
    • Fergusoninidae
    
    
    • Marginidae
    
    
    • Neminidae
    
    
    • Neurochaetidae
    
    
    • Odiniidae
    
    
    • Opomyzidae
    
    
    • Periscelididae
    
    
    • Teratomyzidae
    
    
    • Xenasteiidae
    
    
  
  
  • Superfamily Ephydroidea
    
    • Camillidae
    
    
    • Curtonotidae
    
    
    • Diastatidae
    
    
    • Ephydridae
    
    
    • Drosophilidae
    
    
  
  
  • Superfamily Carnoidea
    
    • Acartophthalmidae
    
    
    • Australimyzidae
    
    
    • Braulidae
    
    
    • Canacidae
    
    
    • Carnidae
    
    
    • Chloropidae
    
    
    • Cryptochaetidae
    
    
    • Inbiomyiidae
    
    
    • Milichiidae
    
    
  
  
  • Superfamily Lonchaeoidea
    
    • Lonchaeidae
    
    
    • Cryptochetidae
    
    
  
  
  • Acalyptratae Incertae Sedis
    
    • Paraleucopidae
  
  

References

External links

• 
  McAlpine, David K. (1958). "A key to the Australian families of Acalptrate Diptera (Insecta)". Records of the Australian Museum. 24 (12): 183–190. doi:10.3853/j.0067-1975.24.1958.650.
  


• 
  "Acalyptratae". Atlas of Living Australia.